Crested wheatgrass, an introduced perennial bunchgrass, has been seeded extensively on the rangelands of western North America. There is a perception that this species is very competitive and that it forms monoculture or low diversity stands where successfully seeded. However, there is limited information on species composition in sites previously seeded to crested wheatgrass. We measured native vegetation and environmental characteristics in areas seeded with crested wheatgrass across the northwestern Great Basin. Plant community composition within these crested wheatgrass stands was variable, from seedings that were near monocultures of crested wheatgrass to those that contained more diverse assemblages of native vegetation, especially shrubs. Environmental factors explained a range of functional group variability from 0% of annual grass density to 56% of large native bunchgrass density. Soil texture appeared to be an important environmental characteristic in explaining vegetation cover and density. Native vegetation was, for all functional groups, positively correlated with soils lower in sand content. Our results suggest environmental differences explain some of the variability of native vegetation in crested wheatgrass stands, and this information will be useful in assessing the potential for native vegetation to co-occupy sites seeded with crested wheatgrass. This research also suggests that crested wheatgrass seedings do not always remain in near monoculture vegetation states as seedings substantially varied in native vegetation composition and abundance with some seeded areas having a more diverse assemblage of native vegetation. In half the sites, there were five or more perennial herbaceous species and 63% of sites contained Wyoming big sagebrush. Although not exclusively true, species most commonly encountered in crested wheatgrass seedings are those that are able to minimize competition with crested wheatgrass via temporal (i.e., Sandberg bluegrass, annual forbs, annual grasses) or spatial (i.e., shrubs) differentiation in resource use. © 2019

Rangelands in the western United States exhibit extremely high temporal variability in seedbed microclimate, and this variability contributes to poor establishment of revegetation species that are typically planted in the fall. We conducted long-term simulations of cumulative germination as a function of planting date and identified alternative germination syndromes based on population-level responses to environmental variability. These germination syndromes reveal ecologically significant differences but also noteworthy similarities in species and seed lot response that can inform rangeland restoration planning and management. Seed germination may occur much sooner than assumed under the traditional paradigm of fall-planting/spring-emergence in the intermountain western United States, and seed germination per se does not appear to be a bottleneck for successful establishment in most years. Instead, simulations of germination response support recent hypotheses that postgermination/preemergent mortality may be the larger contributor to poor seedling establishment. Our data support two general strategies to improve the likelihood of seedling survival into the spring: seeding as late as possible in the fall and active diversification of germination syndromes within a given seed mix. Consistent application of these strategies could increase the probability that some seeds are always available to take advantage of any pulse of seedbed favorability in the late fall, winter, or early spring. © 2019

Dryland ecosystems are particularly vulnerable to erosion generated by livestock grazing. Quantifying this risk across a variety of landscape settings is essential for successful adaptive management, particularly in light of a changing climate. In the Upper Colorado River Basin, there are nearly 25 000 km2 of rangelands with underlying soils derived from Mancos Shale, an erodible and saline geologic parent material. Salinity is a major concern within the Colorado River watershed, much of which is attributed to runoff and leaching from Mancos Shale deposits. In a 60-yr paired-watershed experiment in western Colorado, we used silt fences to measure differences in saline hillslope erosion, including both total sediment yield and concentrations of primary saline constituents (Na and Se), in watersheds that were either exposed to grazing or where livestock was excluded. After accounting for the strong effects of soil type, slope, and antecedent precipitation, we found that grazing increased sediment loss by ≈50% across our 8-yr time series (0.1–1.5 tn ha−1), consistent with levels reported at the watershed scale in early published work from studies at the same location. Eroded sediment Se levels were low and unaffected by grazing history, but Na concentrations were significantly reduced on grazed hillslopes, likely due to depletion of surface Na in soils exposed to chronic soil disturbance by livestock. Variable selection and path analysis identified that biological soil crust (BSC) cover, more than any other variable, explained the differences in sediment yields between grazed and ungrazed watersheds, partially through the enhancement of soil aggregate stability. Our results suggest that BSC cover should be granted heightened consideration in rangeland decision support tools (e.g., state-and-transition models) and that measures to reduce surface disturbance from livestock such as altering the timing or intensity of grazing may be effective for reducing downstream impacts. © 2019

Revegetation of exotic annual grass−invaded rangelands is a primary objective of land managers following wildfires. Controlling invasive annual grasses is essential to increasing revegetation success; however, preemergent herbicides used to control annual grasses prohibit immediate seeding due to nontarget herbicide damage. Thus, seeding is often delayed 1 yr following herbicide application. This delay frequently allows for reinvasion of annual grasses, decreasing the success of revegetation efforts. Incorporating seeds into herbicide protection pods (HPPs) containing activated carbon (AC) permits concurrent high preemergent herbicide application and seeding because AC adsorbs and renders herbicides inactive. While HPPs have, largely in greenhouse studies, facilitated perennial bunchgrass emergence and early growth, their effectiveness in improving establishment of multiple species and functional groups in the field has not been assessed. Five bunchgrass species and two shrub species were seeded at two field sites with high imazapic application rates as bare seed and seed incorporated into HPPs. HPPs significantly improved establishment of sagebrush (Artemesia tridentata Nutt. Spp. wyomingensis Beetle & Young) and crested wheatgrass (Agropyron cristatum [L.] Gaertn.) over the 2-yr study. Three native perennial grass species were protected from herbicide damage by HPPs but had low establishment in both treatments. The two remaining shrub and grass species did not establish sufficiently to determine treatment effects. While establishment of native perennial bunchgrasses was low, this study demonstrates that HPPs can be used to protect seeded bunchgrasses and sagebrush from imazapic, prolonging establishment time in the absence of competition with annual grasses. © 2020

Long-term control of the invasive annual grass cheatgrass is predicated on its biological suppression. Perennial grasses vary in their suppressive ability. We compared the ability of a non-native grass (“Hycrest” crested wheatgrass) and two native grasses (Snake River wheatgrass and bluebunch wheatgrass) to suppress cheatgrass. In a greenhouse in separate tubs, 5 replicates of each perennial grass were established for 96 d, on which two seeds of cheatgrass, 15 cm apart, were then sown in a semicircular pattern at distances of 10 cm, 30 cm, and 80 cm from the established perennial bunchgrasses. Water was not limiting. After 60 d growth, cheatgrass plants were harvested, dried, weight recorded, and tissue C and N quantified. Soil N availability was quantified at each location where cheatgrass was sown, both before sowing and after harvest. Relative to cheatgrass grown at 80 cm, all perennial grasses significantly reduced aboveground biomass at 30 cm (68% average reduction) and at 10 cm (98% average reduction). Sown at 10 cm from established perennial grasses, cheatgrass aboveground biomass was inversely related with perennial grass root mass per unit volume of soil. All cheatgrass sown at 10 cm from “Hycrest” crested wheatgrass died within 38 d. Before sowing of cheatgrass, soil 10 cm from established perennial grasses had significantly less mineral N than soil taken at 30 cm and 80 cm. Relative to cheatgrass tissue N for plants grown at 80 cm, cheatgrass nearest to the established perennial grasses contained significantly less tissue N. All perennial grasses inhibited the NO<inf>2</inf>− to NO<inf>3</inf>− nitrification step; for “Hycrest” crested wheatgrass, soil taken at 10 cm from the plant had a molar proportion of NO<inf>2</inf>− in the NO<inf>2</inf>− + NO<inf>3</inf>− pool of > 90%. In summary, a combination of reduced nitrogen availability, occupation of soil space by perennial roots, and attenuation of the nitrogen cycle all contributed to suppression of cheatgrass. © 2020

A comprehensive understanding of multipaddock, rotational grazing management on rangelands has been slow to develop, and the contribution of adaptive management (Briske et al. 2011) and sufficient scale (Teague and Barnes 2017) have been identified as key omissions. We designed an experiment to compare responses of vegetation and cattle in an adaptively managed, multipaddock, rotational system with that of a season-long, continuous system at scales comparable with those of a working ranch. We hypothesized that 1) year-long rest periods in the adaptively managed, rotational pastures would increase the density and productivity of perennial C<inf>3</inf> graminoids compared with continuously grazed pastures and 2) adaptive management, supported with detailed monitoring data, would result in similar cattle performance in the rotational as in the continuously grazed pastures. However, we found little supporting evidence for grazing management effects on C<inf>3</inf> graminoid abundance or production under either above-average or below-average precipitation conditions during the 5-yr experiment. Furthermore, adaptive rotational grazing resulted in a 12–16% reduction in total cattle weight gain relative to continuous grazing each year. Our work shows that the implementation of adaptive management by a stakeholder group provided with detailed vegetation and animal monitoring data was unable to fully mitigate the adverse consequences of high stock density on animal weight gain. Under adaptive rotational grazing, C<inf>3</inf> perennial grass productivity and stocking rate both increased following above-average precipitation. But when adaptive rotational management was directly compared with continuous grazing with the same increase in stocking rate, continuous grazing achieved similar vegetation outcomes with greater cattle weight gains. We suggest that managers in semiarid rangelands strive to maintain cattle at stock densities low enough to allow for maximal cattle growth rates, while still providing spatiotemporal variability in grazing distribution to enhance rangeland heterogeneity and long-term sustainability of forage production. © 2020

Huisache (Vachellia farnesiana [L.] Wight & Arn.) is a woody species native to the western hemisphere that can invade a variety of native rangeland habitats around the world. In South Texas, it is native yet increasing in density and range while displacing more desirable forage plants, and because it resprouts prolifically it has been difficult to control. We therefore attempted to characterize optimal timing and environmental conditions for herbicide control. We collected weather data in four huisache-invaded pastures along the Texas Coastal Bend and recorded the average huisache phenological stage at those pastures each month. We also analyzed root crown total nonstructural carbohydrate (TNC) and huisache mortality resulting from herbicide treatment each month between April 2012 and November 2014. Huisache mortality and TNC data were analyzed using a randomized complete block analysis of variance, with post hoc tests to separate means; Akaike's information criterion determined best-fit models for mortality data. Root crown TNC increased during May, August, and December, indicating downward translocation and the potential for increased mortality of treated plants. Actual mortality was greatest during the months of May, September, October, and November. The best fit model for mortality was a sixth-order polynomial function of mortality versus month; when month was removed as a model factor, the best fit model was a quadratic function of soil temperature (peaking at ≈24.5°C), combined with a quadratic function of phenology (highest at full canopy stage). These results indicate that huisache is best treated with chemicals when soil temperature is near 24.5°C and during the full canopy stage. In the coastal plains of South Texas these variables coincide with two windows of optimal mortality for huisache treatment: May and September−November. © 2020 The Society for Range Management

Forage and grazing (FG) systems can store a substantial amount of soil organic carbon (SOC) under appropriate land use management and reduce atmospheric CO<inf>2</inf> concentrations. Increasing SOC levels along with many interlinked ecosystem services are essential for increased productivity and sustainability of FG lands (FGLs). Although adoption of improved management practices (MPs) that support SOC sequestration (SOC<inf>q</inf>) is necessary, clear understandings of challenges and opportunities which are sometimes unique to individual FGLs, are also important for implementation of MPs. The objective of this forum paper is to explore the latest scientific knowledge on opportunities to address major challenges for increasing SOC<inf>q</inf> in FGLs. In intensively managed FGLs where the goal is often to maximize yields, lands are heavily fertilized and thus, usually drive towards SOC loss. Diversifications of both forage and grazing species along with strategic grazing plans have been proven as effective MPs for increasing SOC<inf>q</inf>. However, challenge of maintaining productivity levels still remains. Implementing improved grazing for nutrient cycling and integrating forage diversification for increased biodiversity are found to improve soil health attributes, which are critical for SOC<inf>q</inf>. However, to achieve this, we also need to consider site- and soil- specific factors. Extreme climatic events often lead to a decline in soil fertility status, SOC<inf>q</inf> and overall productivity of FGL systems. To address these challenges, uses of models to simulate the FGL systems and have definite choices of suitable MPs are helpful. However, we must be able to access a wide range of datasets to develop system-level adaption strategies that are effective in mitigating these adverse effects. Ultimately, participatory research with novel views and improved perceptions based on the value of SOCq and long-term benefits of the implementation of the best MPs and developing education and outreach materials to enrich the producers’ knowledge gaps are helpful for climate-resilient FGL systems. © 2020 The Society for Range Management

Solarization (covering soil and vegetation with plastic) has long been used in agriculture to control undesirable plants, but solarization of invasive plants in rangelands has shown mixed and species-specific results. Yellow bluestem (Bothriochloa ischaemum (L.) Keng var. songarica (Rupr. ex Fisch & C.A. Mey) Celarier & Harlan), an invasive perennial C<inf>4</inf> grass, is common throughout the southern Great Plains and is not controlled by winter prescribed fire. We tested whether solarization (tarping) with black plastic, combined with winter prescribed fire, could control yellow bluestem. We applied three treatments (with four replicates): solarization (August to November 2017) + fire (January 2018), trimming + fire, and fire only. Results after two growing seasons show that total yellow bluestem cover in solarized + fire plots was reduced to 54% ± 10% (mean ± standard error), lower than trimmed + fire (82% ± 5%, p < 0.01) and fire only plots (78% ± 6%, p = 0.01). Forb cover in solarized + fire plots (15% ± 4%) was much higher than trimmed + fire (4% ± 1%, p < 0.01) and fire only plots (3% ± 1%, p < 0.01). Native forb richness was only slightly higher in solarized + fire plots (16 ± 2 species) compared to fire only (10 ± 2 species, p = 0.08) and trimmed + fire plots (10 ± 1 species, p = 0.08). Interestingly, native forb richness in all plots increased compared to pre-treatment values (2 ± 1 species for all treatments, p < 0.01). Solarization + winter fire can slightly decrease yellow bluestem cover and greatly increase native forb cover, creating islands of diversity in otherwise low-diversity grasslands. However, repeated treatments or alternative techniques will be needed for full control of yellow bluestem. © 2020 The Society for Range Management